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Calibration[ edit ] The molecular clock alone can only say that one time period is twice as long as another:

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Later, the work of Motoo Kimura [6] developed the neutral theory of molecular evolution Clock dating, which predicted a molecular clock.

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This allows fossils to be placed on a branch above an extant organism, rather than being confined to the tips, Clock dating.

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Changing generation times If the Clock dating of new mutations depends at least partly on the number of generations rather than the number of years Population size Genetic drift is stronger in small populations, and so more mutations are effectively neutral Species-specific differences due to differing metabolism, ecology, Clock dating, evolutionary history,

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Molecular and morphological models work together simultaneously, allowing morphology to inform the placement of fossils.

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Molecular clock users have developed workaround solutions using a number of statistical approaches including maximum likelihood techniques and later Bayesian modeling, Clock dating.

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Other sets of species have abundant fossils available, allowing the MCH of constant divergence rates to be tested.

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The molecular clock runs into particular challenges at very short and very long timescales.

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This pattern was also found for all the primate comparisons they tested.

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